Moore and Quick established that the scrotal sac, and its ability to expand or contract, helps to regulate the temperature of the testes. Through this mechanism it enables sperm production and supports the viability of sperm.

Digital facsimile from physiology.org at this link.

(Thanks to Juan Weiss for this reference and its interpretation.)

C. R. Moore, T. F. Gallagher, and F. C. Koch were the first to obtain a potent testicular extract containing the male sex hormone, androsterone, later obtained in crystalline form by Butenandt. They also gave a detailed account of the capon-comb test for the assay of the male hormone.